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and prefer to work with a small number of parameters rather than many. These advantages should apply to the expanded social fitness model that includes and distinguishes kin, kith, and kind.

ACKNOWLEDGMENTS

I thank Francisco Ayala and John Avise for helping to organize the National Academy of Sciences Sackler symposium on cooperation and conflict. For comments on the manuscript, I thank two anonymous referees, Joan Strassmann, Michael Whitlock, Stuart West, Kevin Foster, Claire El Mouden, and the Oxford Social Evolution Group. Our research is supported by U.S. National Science Foundation Grants DEB 0816690 and DEB 0918931.



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