Living fish and fossil fish share many similarities, but the fossil fish clearly belongs to a different species that no longer exists. The progression of species found in the fossil record provides powerful evidence for evolution.

with characteristics enabling them to occupy ecological niches not occupied by similar organisms have a greater chance of surviving. Over time—as the next chapter discusses in more detail—species have diversified and have occupied more and more ecological niches to take advantage of new resources.

Evolution explains something else as well. During the billions of years that life has been on earth, it has played an increasingly important role in altering the planet's physical environment. For example, the composition of our atmosphere is partly a consequence of living systems. During photosynthesis, which is a product of evolution, green plants absorb carbon dioxide and water, produce organic compounds, and release oxygen. This process has created and continues to maintain an atmosphere rich in oxygen. Living communities also profoundly affect weather and the movement of water among the oceans, atmosphere, and land. Much of the rainfall in the forests of the western Amazon basin consists of water that has already made one or more recent trips through a living plant. In addition, plants and soil microorganisms exert important controls over global temperature by absorbing or emitting ''greenhouse gases" (such as carbon dioxide and methane) that increase the earth's capacity to retain heat.

In short, biological evolution accounts for three of the most fundamental features of the world around us: the similarities among living things, the diversity of life, and many features of the physical world we inhabit. Explanations of these phenomena in terms of evolution draw on results from physics, chemistry, geology, many areas of biology, and other sciences. Thus, evolution is the central organizing principle that biologists use to understand the world. To teach biology without explaining evolution deprives students of a powerful concept that brings great order and coherence to our understanding of life.

The teaching of evolution also has great practical value for students. Directly or indirectly, evolutionary biology has made many contributions to society. Evolution explains why many human pathogens have been developing resistance to formerly effective drugs and suggests ways of confronting this increasingly serious problem (this issue is discussed in greater detail in Chapter 2). Evolutionary biology has also

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